Scotland’s most experienced university principal has called for a review of the funding and shape of higher education, warning against “stumbling from year to year”.

Sir Anton Muscatelli, who retires this month after 16 years as the principal and vice-chancellor of Glasgow University, says universities need a plan which could see major changes in the courses they teach and the research they carry out.

He told the BBC that charging tuition fees to Scottish students is unlikely to secure support at Holyrood so Scotland needs to work out what it wants from higher education and then decide how to pay for it.

His call comes ahead of two crucial reports, due to be published on Friday by the Scottish Funding Council (SFC), about the financial position of universities and of further education colleges.

Both studies have been delayed since early this year – and should reflect the latest financial figures up to June.

This comes as analysis seen by the BBC suggests Scottish government finance for universities and further education colleges has been squeezed in real terms in the past six years by about a fifth.

FE colleges say they are having to turn away qualified applicants and that apprenticeship course are over-subscribed, while universities are limiting the number of places they can offer Scottish students.

In recent years, some have subsidised the cost of educating Scottish students by sharply increasing their recruitment of foreign students but those numbers have started to decline.

Earlier this year, Dundee University was hit by a severe financial crisis.

A report into the near-collapse of the university said it had continued with increased spending despite a drop in foreign student fee income.

Its principal went before a Scottish Parliament committee and admitted incompetence and senior board members resigned over the mismanagement.

Dundee has so far required a Scottish government bailout of more than £40m.

It is an extreme example but other Scottish universities are also under financial strain.

Edinburgh University is planning cuts of £140m, including job losses.

Its principal, Sir Peter Matheson, has called for a “radical re-wiring” to respond to the funding challenge.

Both institutions and others have seen recent strikes by academic staff to highlight the effect of cuts.

Students at Scottish universities with permanent homes in Scotland don’t pay tuition fees.

Instead, the universities get money from the government for a set number of Scottish students each year.

However, they claim that the amount they receive does not cover their costs – and in most cases is not as much as the £9,500 a year that students from the rest of the UK usually pay.

Foreign students in Scotland pay much higher fees – in many cases nearly three times more than UK students.

Most of them come to the UK to do post-graduate Masters degree courses.

This has been an area that has provided a welcome source of funding in recent years but it has fallen away sharply for several reasons, including new UK immigration rules and competition from other countries.

There is no sign of Holyrood ministers or their opponents moving to end ‘free tuition’ for Scottish students, so they need to find other ways of tackling their financial problems.

The BBC has had access to a new study carried out by David Bell, a professor of economics at Stirling University and expert in public finance.

The study, written for the Royal Society of Edinburgh, details how funding per Scottish university student has fallen over the past five years due to the effect of inflation.

In real terms, the funding shrunk by 22% between 2019-20 and 2023-24, Prof Bell says.

The academic also used publicly available data to show that the fees universities receive fall far short of what they estimate are the costs of educating students.

For instance, in modern languages, the £7,421 annual funding per student is half of what they say is required.

In dentistry, the Scottish government last year paid £19,580 per student.

This is reckoned to be at least £9,000 short of the cost to universities.

The funding for veterinary studies, law, social work, media studies, drama, architecture, planning, anatomy, chemistry, mineral engineering, philosophy and religion would have to be increased by at least 70% to reach the actual cost of educating the average student, Prof Bell says.

Universities also say their research funding, from governments and other sources, fails to cover costs of the overheads they have in facilities.

The number of foreign students at Scottish universities has nearly doubled since 2006-07, when it was 24,200.

The figure reached a peak of 47,700 in 2022-23 before falling back.

Prof Bell’s report details the ways in which the money from overseas students is threatened by changes to immigration requirements.

There are new rules that bar students from bringing their families, as well as a reduction in the time foreign nationals are allowed to work in the UK after graduating.

Some universities are much more exposed than others to volatile foreign student income.

The University of the West of Scotland and Glasgow Caledonian have three quarters of the places on their taught post-grad courses filled by non-UK students.

Older universities such as Glasgow and Edinburgh have many more such students, and more than half of whom are from overseas.

The number of places for Scottish students doing their first degree is capped in each university, due to constraints in Scottish government funding.

But there is no limit to the number of places that can be taken by students from the rest of the UK.

They pay the same level of annual tuition fees that they would pay in the rest of the UK.

For the new academic year, that is £9,535, a rise of of 3.1%.

Universities across the UK say this amount has not increased by very much for more than a decade and claim that it also falls short of the actual cost of educating students.

Universities such as Edinburgh and St Andrews have almost as many students from England, Wales and Northern Ireland as they do from Scotland, with roughly a third from each.

Glasgow, by contrast, is 66% Scottish and 15% are from the rest of the UK.

For newer universities, with less of a reputation and prestige, income from the rest of the UK students is more limited.

At Glasgow Caledonian and the University of the Highlands and Islands, the Scottish share is more than 95%.

One effect of the Scottish government providing tuition fees for students from Scotland is that graduates themselves leave university with much less debt.

The study cites research that indicates the average Scot graduated in 2023 with debt of £15,430, while the figure for the rest of the UK was £44,940.

Prof Bell’s research gives extensive detail on earnings five years after they graduate.

His figures, based on the 2021-22 tax year, vary widely.

They show a computing graduate from St Andrews was earning an average £77,000, three times as much as one from the University of the West of Scotland.

The lowest pay was for a typical graduate in art and design from the University of the Highlands and Islands, on £15,300.

Other high-earning graduates were in medicine, dentistry and economics.

Among lower earners were those with degrees in media, sociology and performing arts.

The report says that the annual earnings of those with qualifications in health-related and STEM subjects tend to exceed the Scottish average, while those with arts qualifications tend to earn less.

Prof Bell points out that funding is mainly allocated to universities based on the previous year’s funds.

He argues that this gives little scope for innovation and change.

Sir Anton Muscatelli also talks about the case for universities becoming more different from each other.

The Glasgow University principal says there will be a need for more of them to share resources and show they are operating efficiently, with mergers being an option.

Looking to next year’s Holyrood election, he says politicians need to acknowledge that the education sector is going to be crucial to the success of the Scottish economy.

He said: “It really is imperative that we don’t stumble from year to year [but] think about how this is going to be shaped or resourced.

“We need a strategic plan as a country, which is why I suspect after 2026, there will need to be a look at the shape and size of the Scottish [university] sector.

“I hope this is done in a rational way. I strongly believe in a publicly-funded sector, I hope that can be achieved, but let’s start with the question: what sector does Scotland need?”

The government is looking at ways to financially support the companies in Jaguar Land Rover’s (JLR) supply chain.

JLR halted car production at the end of August after a cyber attack forced it to shut down its IT networks. Its factories remain suspended until next month at the earliest.

Fears are growing that some suppliers, in particular the smaller firms who solely rely on JLR’s business, could go bust without support.

One idea being explored is the government buying the component parts the suppliers build, to keep them in business until JLR’s production lines are up and running again.

Unions had called for a Covid-style furlough scheme, but ministers have ruled this out given its likely cost, sources have told the BBC.

Another option being considered is providing government-backed loans to suppliers, though this is understood to be out unpopular with suppliers.

The purchase and stockpiling of car parts by the government is also an option on the table, but this would present considerable logistical challenges.

JLR’s manufacturing process relies on the right part arriving at the right place, at the right time.

However, industry experts agree doing nothing risks firms in the supply chain, which employs tens of thousands of workers, facing bankruptcy.

The Business and Trade Select Committee is due to meet on Thursday afternoon to hear testimonies from businesses in JLR’s supply chain because of deep concern for some of these businesses to remain viable.

This evidence will be shared with the government afterwards.

Senior government figures are concerned about a pattern of cyber attacks on UK institutions and businesses, such as the British Library, Marks & Spencer, and the Co-op.

A group calling itself Scattered Lapsus$ Hunters has claimed responsibility for the hack on JLR, Marks & Spencer, and Co-op.

An investigation is under way into the cyber attack on JLR, which is believed to be costing the company at least £50m a week in lost production.

JLR would normally expect to build more than 1,000 cars a day at its three factories in Solihull and Wolverhampton in West Midlands and Halewood in Merseyside.

However, workers were sent home following the hack – which first came to light on 1 September – with no firm return date.

About 30,000 people are directly employed at those plants with a further 100,000 working in the firm’s supply chain.

On Tuesday, the business secretary and industry minister visited the West Midlands for the first time since the incident to meet JLR and the firms in its supply chain.

Additional reporting by Pritti Mistry